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Fossil gibbons are exceedingly rare, with much of the hylobatid fossil record and, consequently, hylobatid evolutionary history remaining unknown. Kapi ramnagarensis was described as a stem hylobatid on the basis of an isolated lower right M3 from ~13.0-12.5 Ma deposits surrounding Ramnagar (J&K), India. This interpretation was recently challenged, with alternative hypotheses suggesting that it is instead a stem catarrhine or a strangely derived pliopithecoid that has converged on hylobatid morphology. A series of morphological features were said to distinguish Kapi from fossil and extant hylobatids; notably, however, none of these features were examined or compared using quantitative analyses. Here, we further examine the dental morphology of Kapi, providing quantitative analyses to critically evaluate the hypothesis that Kapi represents a stem catarrhine or pliopithecoid rather than a stem hylobatid. Results demonstrate that none of the claimed differences between Kapi and hylobatids hold up under closer scrutiny, and multivariate discriminant analyses taking size and shape into account strongly support Kapi as a hylobatid with high posterior probabilities. Although only represented by a single lower molar, Kapi remains the most compelling candidate for the earliest known hylobatid in the fossil record and thus likely documents the simultaneous arrival of lesser and great apes to Asia during the Middle Miocene. 

The fossil record of ‘lesser apes’ (i.e. hylobatids = gibbons and siamangs) is virtually non-existent before the latest Miocene of East Asia. However, molecular data strongly and consistently suggest that hylobatids should be present by approximately 20 Ma; thus, there are large temporal, geographical, and morphological gaps between early fossil apes in Africa and the earliest fossil hylobatids in China. Here, we describe a new approximately 12.5–13.8 Ma fossil ape from the Lower Siwaliks of Ramnagar, India, that fills in these long-standing gaps with implications for hylobatid origins. This ape represents the first new hominoid species discovered at Ramnagar in nearly a century, the first new Siwalik ape taxon in more than 30 years, and likely extends the hylobatid fossil record by approximately 5 Myr, providing a minimum age for hylobatid dispersal coeval to that of great apes. The presence of crown hylobatid molar features in the new species indicates an adaptive shift to a more frugivorous diet during the Middle Miocene, consistent with other proposed adaptations to frugivory (e.g. uricase gene silencing) during this time period as well. 

Abstract Over the last several decades, the study of Earth surface processes has progressed from a descriptive science to an increasingly quantitative one due to advances in theoretical, experimental, and computational geosciences. The importance of geomorphic forecasts has never been greater, as technological development and global climate change threaten to reshape the landscapes that support human societies and natural ecosystems. Here we explore best practices for developing socially relevant forecasts of Earth surface change, a goal we are calling “earthcasting”. We suggest that earthcasts have the following features: they focus on temporal (∼1–∼100 years) and spatial (∼1 m–∼10 km) scales relevant to planning; they are designed with direct involvement of stakeholders and public beneficiaries through the evaluation of the socioeconomic impacts of geomorphic processes; and they generate forecasts that are clearly stated, testable, and include quantitative uncertainties. Earthcasts bridge the gap between Earth surface researchers and decision‐makers, stakeholders, researchers from other disciplines, and the general public. We investigate the defining features of earthcasts and evaluate some specific examples. This paper builds on previous studies of prediction in geomorphology by recommending a roadmap for (a) generating earthcasts, especially those based on modeling; (b) transforming a subset of geomorphic research into earthcasts; and (c) communicating earthcasts beyond the geomorphology research community. Earthcasting exemplifies the social benefit of geomorphology research, and it calls for renewed research efforts toward further understanding the limits of predictability of Earth surface systems and processes, and the uncertainties associated with modeling geomorphic processes and their impacts.